Stromatolites are not only Earth's oldest of fossils, but are intriguing in that they are our singular visual portal into deep time on earth, the emergence of life, and the evolving of the beautiful forms of life of modern time. A small piece of stromatolites encodes biological activity perhaps spanning thousands of years. In broad terms, stromatolites are fossil evidence of the prokaryotic life that remains today, as it has always been, the preponderance of biomass in the biosphere. For those that subscribe
Scientists disagree on how to define stromatolites. A common definition goes something like: A lamiated rock formed by the growth of blue-green algae (i.e., cyanobacteria)". This definition is, in fact, such a gross oversimplification as be scientifically useless. It does contain a modicum of truth, however, in that the largest volume of stromatolite formations was likely formed by biogenic processes involving photosynthetic cyanobacteria. Cyanobacteria’s metabolic byproduct, oxygen, rusted the earth, pumped enormous oxygen poison to them into earth’s atmosphere, and in so doing paved the way for aerobic-based life to emerge and diversify; cyanobacteria’s contributions to life led to their own prodigious decline.
Stromatolites                  and their close cousins the thrombolites, are rock-like buildups                  of microbial mats that form in limestone- or dolostone-forming                  environments. Together with oncoids (formerly called "algal                  biscuits" or "Girvanella"), they typically form                  by the baffling, trapping, and precipitation of particles by communities                  of microorganisms such as bacteria and algae. In some cases, they                  can form inorganically, for example when seawaters are oversaturated                  with certains chemicals resulting in precipitation. Stromatolites                  are defined as laminated accretionary structures that have synoptic                  relief (i.e., they stick up above the seafloor). Stromatolite-building                  communities include the oldest known fossils,                  dating back some 3.5 billion years when the environments of Earth                  were too hostile to support life as we know it today. We can presume                  that the microbial communities consisted of complex consortia                  of species with diverse metabolic needs, and that competition                  for resources and differing motility among them created the intricate                  structures we observe in these ancient fossils. Microbial communities                  diversified through time, with eukaryotic organisms eventually                  joining the mix.
Excluding some exceedingly rare Precambrian fossils such as the Ediacaran fauna, stromatolites and are the only fossils encoding the first 7/8th of the history of life on earth. They encode the role that ancient microorganisms played in the evolution of life on earth and in shaping earth's environments. The fossil record of stromatolites is astonishingly extensive, spanning some four billion years of geological history with the forming organisms possibly having occupied every conceivable environment that ever existed on earth. Today, stromatolites are nearly extinct in marine environments, living a precarious existence in only a few localities worldwide. Modern stromatolites were first discovered in Shark Bay, Australia in 1956, and throughout western Australia in both marine and non-marine environments. New stromatolite localities have continued to be discovered in various places such as the Bahamas, the Indian Ocean and Yellowstone National Park, to name but a few localities.








 
